Energy & Ecosystems (A Level Only)

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Primary Producers

  • Living organisms need organic compounds such as carbohydrates, proteins and lipids to respire and build tissues.
  • In ecosystems, organic compounds are synthesised by plants during photosynthesis.
  • Plants use carbon from carbon dioxide to produce organic molecules: CO₂ from the atmosphere in terrestrial ecosystems, and dissolved CO₂ in aquatic ecosystems.
  • Plants are primary producers because they produce their own organic molecules.
  • These organic compounds are passed to other organisms when herbivores consume plant tissue.

Photosynthetic Products

  • Plants synthesise sugars during photosynthesis.
  • Most sugars are used by the plant as respiratory substrates.
  • Remaining sugars are used to make other biological molecules that form plant biomass.
  • Starch is a complex carbohydrate that functions as an energy storage molecule inside plant cells.
  • Cellulose is a complex carbohydrate that forms a structural component of plant cell walls.
  • Lipids are made from sugars and are used for energy storage and formation of the waxy cuticle on leaves.
  • Proteins are made by combining sugars with nitrates to make amino acids, e.g. membrane proteins and enzymes.

Biomass

  • Biomass is the mass of living material present in an organism or tissue sample.
  • Biomass can be measured as the dry mass per given area (mass of tissue after water has been removed).
  • Dry mass is more reliable than wet mass because the water content of living tissue can vary.
  • Units for dry mass are mass per unit area, e.g. kg m⁻².
  • Biomass can also be measured as the mass of carbon per given area, taken to be 50% of the dry mass.
  • Dry mass of a small sample can be scaled up to calculate the biomass of a total population or area.

Measuring Dry Mass and Stored Energy

  • To find dry mass, dry the sample until it contains no more water, using a crucible, oven and digital balance.
  • Set the oven to a low temperature so the sample does not burn and lose biomass.
  • Weigh the crucible and sample at frequent intervals until the mass stops decreasing (fully dehydrated), then subtract the original crucible mass.
  • Limitations: drying can take several days for larger samples, and very precise equipment may not be available.
  • Calorimetry can estimate the chemical energy stored in dry biomass.
  • In a simple calorimeter, burn the sample beneath a beaker of water and calculate: energy per gram = (mass of water × temperature change × 4.2) ÷ mass of sample.
  • Limitations: heat may be transferred to the surroundings rather than the water; a bomb calorimeter is more accurate but expensive.

Gross Primary Production

  • Gross primary production (GPP) is the chemical energy stored in plant biomass, in a given area or volume.
  • In terrestrial plants, GPP is expressed per unit area, e.g. J m⁻² or g m⁻².
  • In aquatic environments, GPP is expressed per unit volume, e.g. kJ m⁻³ or kg m⁻³.
  • Biomass = stored chemical energy, so a measure of mass can represent energy.
  • GPP is not a rate, so its units do not include time; units including time measure productivity.

Net Primary Production

  • Net primary production (NPP) is the chemical energy stored in plant biomass after respiratory losses to the environment have been taken into account.
  • Plants convert light energy to chemical energy in glucose during photosynthesis; the total stored is GPP.
  • Some energy stored during GPP is lost to the environment, e.g. as waste heat generated during respiration.
  • NPP is calculated as NPP = GPP − R, where R = respiratory loss to the environment.
  • NPP is expressed as energy per unit area or volume, e.g. J m⁻² or J m⁻³.
  • NPP represents the energy available to consumers in the ecosystem.
  • The NPP equation applies only to producers, so it includes only respiratory loss, not losses in faeces and urine.

Trophic Levels and Net Production of Consumers

  • NPP is available for plant growth and reproduction, and for transfer to consumers when plant biomass is eaten by herbivores or broken down by decomposers.
  • Consumers gain energy from the tissues of other organisms.
  • Ingested chemical energy is either transferred to the consumer's tissues or lost to the environment.
  • The energy transferred to consumer tissues is the net production of consumers (also called secondary production).
  • Net production of consumers is calculated as N = I − (F + R), where I = chemical energy in ingested food, F = energy lost in faeces and urine, R = respiratory losses.
  • Energy is lost in faeces because consumers cannot digest all food (e.g. cellulose, fur and bones).
  • Energy is lost in urine because excess amino acids are converted into urea for excretion.
  • Energy is lost during respiration as heat, which is radiated into the environment.

Primary & Secondary Productivity

  • Productivity is the rate of primary or secondary production.
  • Primary production = transfer of energy to the tissues of plants.
  • Secondary production = transfer of energy to the tissues of consumers (net production of consumers).
  • Productivity is expressed as mass or energy per unit area or volume over a given time period, e.g. g m⁻² day⁻¹, J m⁻² day⁻¹, kJ ha⁻¹ year⁻¹.
  • Production has no unit of time; productivity is a rate and must include time.
  • In aquatic environments, productivity may be measured per unit volume, e.g. g m⁻³ yr⁻¹.

Calculating Productivity & Efficiency

  • Net productivity of producers uses NPP = GPP − R; productivity is a rate, so units include time.
  • Net productivity of consumers uses N = I − (F + R).
  • Efficiency of energy transfer is the percentage of energy transferred between trophic levels.
  • % efficiency = (productivity of trophic level ÷ energy available from level below) × 100.
  • Some calculations require multiple steps, e.g. first calculate net productivity, then use it in the efficiency equation.
  • Always check the units to decide whether the value given is productivity (a rate) or production.

Farming & Production

  • Farming practices can increase production to increase food available for human consumption by simplifying food webs and reducing respiratory loss.
  • Food webs can be simplified by removing pests from crops using chemical pesticides or biological pest control.
  • Pests reduce crop biomass and cause crop plants to expend energy on herbivory defences rather than growth, reducing NPP.
  • Respiratory loss in livestock can be reduced by restricting movement (keeping animals in pens) to reduce energy for muscle activity.
  • Keeping animals warm in heated indoor environments reduces energy used for temperature regulation.
  • Antibiotics may be given to healthy animals to prevent infection, reducing energy used by the immune system.
  • These practices give higher energy outputs in less time at lower cost, but raise ethical concerns about animal welfare.
  • Percentage yield = (actual yield ÷ theoretical yield) × 100; yield is below 100% due to factors such as non-ideal temperature, light intensity, CO₂ concentration, pests or disease.

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Soal latihan

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  1. 1.Which term describes plants in an ecosystem because they produce their own organic molecules?

    Easy
    • APrimary producers
    • BPrimary consumers
    • CDecomposers
    • DSecondary consumers
  2. 2.Which of the following is the best definition of biomass?

    Easy
    • AThe mass of living material present in an organism or tissue sample
    • BThe total number of organisms in a given area
    • CThe energy stored in an organism's tissues
    • DThe dry mass of an organism per unit area
  3. 3.Why is dry biomass a more accurate measurement than wet biomass?

    Easy
    • AThe water content of living tissue can vary
    • BDry biomass includes the mass of water
    • CWet biomass is measured in different units
    • DDry biomass measures the number of organisms
  4. 4.Which of the following is the correct definition of gross primary production (GPP)?

    Medium
    • AThe chemical energy stored in plant biomass, in a given area or volume
    • BThe chemical energy stored in plant biomass after respiratory losses have been taken into account
    • CThe rate of energy transfer to plant tissues
    • DThe total energy ingested by consumers in a food chain
  5. 5.Which equation correctly represents net primary production (NPP)?

    Medium
    • ANPP = GPP − R
    • BNPP = GPP + R
    • CNPP = I − (F + R)
    • DNPP = GPP × R
  6. 6.In the equation N = I − (F + R) for consumers, what does F represent?

    Medium
    • AThe chemical energy lost to the environment in faeces and urine
    • BThe chemical energy in ingested food
    • CThe respiratory losses to the environment
    • DThe net production of the consumer
  7. 7.Which of the following units is appropriate for gross primary production in a terrestrial ecosystem?

    Easy
    • AkJ m⁻²
    • BkJ m⁻² yr⁻¹
    • CkJ m⁻³
    • DkJ m⁻³ yr⁻¹
  8. 8.Which of the following units indicates a measure of productivity rather than production?

    Easy
    • AkJ m⁻² yr⁻¹
    • BkJ m⁻²
    • Cg m⁻²
    • Dkg m⁻³

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