Energy & Ecosystems (A Level Only)
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Primary Producers
- Living organisms need organic compounds such as carbohydrates, proteins and lipids to respire and build tissues.
- In ecosystems, organic compounds are synthesised by plants during photosynthesis.
- Plants use carbon from carbon dioxide to produce organic molecules: CO₂ from the atmosphere in terrestrial ecosystems, and dissolved CO₂ in aquatic ecosystems.
- Plants are primary producers because they produce their own organic molecules.
- These organic compounds are passed to other organisms when herbivores consume plant tissue.
Photosynthetic Products
- Plants synthesise sugars during photosynthesis.
- Most sugars are used by the plant as respiratory substrates.
- Remaining sugars are used to make other biological molecules that form plant biomass.
- Starch is a complex carbohydrate that functions as an energy storage molecule inside plant cells.
- Cellulose is a complex carbohydrate that forms a structural component of plant cell walls.
- Lipids are made from sugars and are used for energy storage and formation of the waxy cuticle on leaves.
- Proteins are made by combining sugars with nitrates to make amino acids, e.g. membrane proteins and enzymes.
Biomass
- Biomass is the mass of living material present in an organism or tissue sample.
- Biomass can be measured as the dry mass per given area (mass of tissue after water has been removed).
- Dry mass is more reliable than wet mass because the water content of living tissue can vary.
- Units for dry mass are mass per unit area, e.g. kg m⁻².
- Biomass can also be measured as the mass of carbon per given area, taken to be 50% of the dry mass.
- Dry mass of a small sample can be scaled up to calculate the biomass of a total population or area.
Measuring Dry Mass and Stored Energy
- To find dry mass, dry the sample until it contains no more water, using a crucible, oven and digital balance.
- Set the oven to a low temperature so the sample does not burn and lose biomass.
- Weigh the crucible and sample at frequent intervals until the mass stops decreasing (fully dehydrated), then subtract the original crucible mass.
- Limitations: drying can take several days for larger samples, and very precise equipment may not be available.
- Calorimetry can estimate the chemical energy stored in dry biomass.
- In a simple calorimeter, burn the sample beneath a beaker of water and calculate: energy per gram = (mass of water × temperature change × 4.2) ÷ mass of sample.
- Limitations: heat may be transferred to the surroundings rather than the water; a bomb calorimeter is more accurate but expensive.
Gross Primary Production
- Gross primary production (GPP) is the chemical energy stored in plant biomass, in a given area or volume.
- In terrestrial plants, GPP is expressed per unit area, e.g. J m⁻² or g m⁻².
- In aquatic environments, GPP is expressed per unit volume, e.g. kJ m⁻³ or kg m⁻³.
- Biomass = stored chemical energy, so a measure of mass can represent energy.
- GPP is not a rate, so its units do not include time; units including time measure productivity.
Net Primary Production
- Net primary production (NPP) is the chemical energy stored in plant biomass after respiratory losses to the environment have been taken into account.
- Plants convert light energy to chemical energy in glucose during photosynthesis; the total stored is GPP.
- Some energy stored during GPP is lost to the environment, e.g. as waste heat generated during respiration.
- NPP is calculated as NPP = GPP − R, where R = respiratory loss to the environment.
- NPP is expressed as energy per unit area or volume, e.g. J m⁻² or J m⁻³.
- NPP represents the energy available to consumers in the ecosystem.
- The NPP equation applies only to producers, so it includes only respiratory loss, not losses in faeces and urine.
Trophic Levels and Net Production of Consumers
- NPP is available for plant growth and reproduction, and for transfer to consumers when plant biomass is eaten by herbivores or broken down by decomposers.
- Consumers gain energy from the tissues of other organisms.
- Ingested chemical energy is either transferred to the consumer's tissues or lost to the environment.
- The energy transferred to consumer tissues is the net production of consumers (also called secondary production).
- Net production of consumers is calculated as N = I − (F + R), where I = chemical energy in ingested food, F = energy lost in faeces and urine, R = respiratory losses.
- Energy is lost in faeces because consumers cannot digest all food (e.g. cellulose, fur and bones).
- Energy is lost in urine because excess amino acids are converted into urea for excretion.
- Energy is lost during respiration as heat, which is radiated into the environment.
Primary & Secondary Productivity
- Productivity is the rate of primary or secondary production.
- Primary production = transfer of energy to the tissues of plants.
- Secondary production = transfer of energy to the tissues of consumers (net production of consumers).
- Productivity is expressed as mass or energy per unit area or volume over a given time period, e.g. g m⁻² day⁻¹, J m⁻² day⁻¹, kJ ha⁻¹ year⁻¹.
- Production has no unit of time; productivity is a rate and must include time.
- In aquatic environments, productivity may be measured per unit volume, e.g. g m⁻³ yr⁻¹.
Calculating Productivity & Efficiency
- Net productivity of producers uses NPP = GPP − R; productivity is a rate, so units include time.
- Net productivity of consumers uses N = I − (F + R).
- Efficiency of energy transfer is the percentage of energy transferred between trophic levels.
- % efficiency = (productivity of trophic level ÷ energy available from level below) × 100.
- Some calculations require multiple steps, e.g. first calculate net productivity, then use it in the efficiency equation.
- Always check the units to decide whether the value given is productivity (a rate) or production.
Farming & Production
- Farming practices can increase production to increase food available for human consumption by simplifying food webs and reducing respiratory loss.
- Food webs can be simplified by removing pests from crops using chemical pesticides or biological pest control.
- Pests reduce crop biomass and cause crop plants to expend energy on herbivory defences rather than growth, reducing NPP.
- Respiratory loss in livestock can be reduced by restricting movement (keeping animals in pens) to reduce energy for muscle activity.
- Keeping animals warm in heated indoor environments reduces energy used for temperature regulation.
- Antibiotics may be given to healthy animals to prevent infection, reducing energy used by the immune system.
- These practices give higher energy outputs in less time at lower cost, but raise ethical concerns about animal welfare.
- Percentage yield = (actual yield ÷ theoretical yield) × 100; yield is below 100% due to factors such as non-ideal temperature, light intensity, CO₂ concentration, pests or disease.
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Câu hỏi luyện tập
Xem trước miễn phí — 8 trên 64 câu hỏi. Đăng ký để xem tất cả.
1.Which term describes plants in an ecosystem because they produce their own organic molecules?
Easy- APrimary producers
- BPrimary consumers
- CDecomposers
- DSecondary consumers
2.Which of the following is the best definition of biomass?
Easy- AThe mass of living material present in an organism or tissue sample
- BThe total number of organisms in a given area
- CThe energy stored in an organism's tissues
- DThe dry mass of an organism per unit area
3.Why is dry biomass a more accurate measurement than wet biomass?
Easy- AThe water content of living tissue can vary
- BDry biomass includes the mass of water
- CWet biomass is measured in different units
- DDry biomass measures the number of organisms
4.Which of the following is the correct definition of gross primary production (GPP)?
Medium- AThe chemical energy stored in plant biomass, in a given area or volume
- BThe chemical energy stored in plant biomass after respiratory losses have been taken into account
- CThe rate of energy transfer to plant tissues
- DThe total energy ingested by consumers in a food chain
5.Which equation correctly represents net primary production (NPP)?
Medium- ANPP = GPP − R
- BNPP = GPP + R
- CNPP = I − (F + R)
- DNPP = GPP × R
6.In the equation N = I − (F + R) for consumers, what does F represent?
Medium- AThe chemical energy lost to the environment in faeces and urine
- BThe chemical energy in ingested food
- CThe respiratory losses to the environment
- DThe net production of the consumer
7.Which of the following units is appropriate for gross primary production in a terrestrial ecosystem?
Easy- AkJ m⁻²
- BkJ m⁻² yr⁻¹
- CkJ m⁻³
- DkJ m⁻³ yr⁻¹
8.Which of the following units indicates a measure of productivity rather than production?
Easy- AkJ m⁻² yr⁻¹
- BkJ m⁻²
- Cg m⁻²
- Dkg m⁻³
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